Vocal cues

Imprinting is an important type of behavior by which an animal learns specific concepts and identifies certain objects or individuals that are essential for survival. Imprinting events almost always occur very early in the life of an animal, during critical periods or time frames when the animal is most sensitive to environmental cues and influences. The phenomenon occurs in a variety of species, but it is most pronounced in the homeothermic (warmblooded) and socially oriented higher vertebrate species, especially mammals and birds.

Imprinting is learned behavior. Most learned behavior falls within the domain of exogenous behavior, or behavior that an animal obtains by its experiences with fellow conspecifics (members of the same species) and the environment. Imprinting, however, is predominantly, if not exclusively, an endogenous behavior, which is a behavior that is genetically encoded within the individual. An individual is born with the capacity to imprint. The animal's cellular biochemistry and physiology will determine when in its development it will imprint. The only environmental influence of any consequence in imprinting is the object of the imprint during the critical period. Ethologists, scientists who study animal behavior, debate the extent of endogenous and exogenous influences upon animal behavior. Most behaviors involve a combination of both, although one type may be more pronounced than the other.

The capacity for an animal to imprint is genetically determined and, therefore, is inherited. This type of behavior is to the animal's advantage for critical situations that must be correctly handled the first time they occur. Such behaviors include the identification of one's parents (especially one's mother), the ability to navigate, the ability to identify danger, and even the tendency to perform the language of one's own species. Imprinting behaviors generally are of high survival value and hence must be programmed into the individual via the genes. Biological research has failed to identify many of the genes that are responsible for imprinting behaviors, although the hormonal basis of imprinting is well understood. Most imprinting studies have focused upon the environmental signals and developmental state of the individual during the occurrence of imprinting.

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